"Unsigned parameter: email" when it is sent to the "Evasion" subscription

I have a simple Ruby on Rails application with Devise for authentication. I would like users to register using a username instead of email and try to implement this as described here: Invent authentication with a username instead of email

The twist is that I still want every user to have an email address and require it during registration. So, I left the "email" field in the registration form.

The problem is that when I fill out the registration form using a valid email address (or any other email value), Devise reports the error: "Email cannot be empty."

Why is email considered blank and how can I fix it?

class ApplicationController < ActionController::Base
  # Prevent CSRF attacks by raising an exception.
  # For APIs, you may want to use :null_session instead.
  protect_from_forgery with: :exception

  before_action :configure_permitted_parameters, if: :devise_controller?

  protected

  def configure_permitted_parameters
    devise_parameter_sanitizer.for(:sign_up) { |u| u.permit(:username, :email, :password, :password_confirmation, :remember_me) }
    devise_parameter_sanitizer.for(:sign_in) { |u| u.permit(:username, :password, :remember_me) }
    devise_parameter_sanitizer.for(:account_update) { |u| u.permit(:username, :email, :password, :password_confirmation, :current_password) }
  end
end

User model

class User < ActiveRecord::Base
  # Include default devise modules. Others available are:
  # :confirmable, :lockable, :timeoutable and :omniauthable
  devise :database_authenticatable, :registerable,
         :recoverable, :rememberable, :trackable, :validatable
end

Type for registration

<h2>Sign up</h2>

<%= form_for(resource, as: resource_name, url: registration_path(resource_name)) do |f| %>
  <%= devise_error_messages! %>

  <div class="field">
    <%= f.label :email %><br />
    <%= f.email_field :email, autofocus: true %>
  </div>

  <div class="field">
    <%= f.label :username %><br />
    <%= f.text_field :username %>
  </div>

  <div class="field">
    <%= f.label :password %>
    <% if @validatable %>
    <em>(<%= @minimum_password_length %> characters minimum)</em>
    <% end %><br />
    <%= f.password_field :password, autocomplete: "off" %>
  </div>

  <div class="field">
    <%= f.label :password_confirmation %><br />
    <%= f.password_field :password_confirmation, autocomplete: "off" %>
  </div>

  <div class="actions">
    <%= f.submit "Sign up" %>
  </div>
<% end %>

<%= render "devise/shared/links" %>

devise.rb modified to include:

config.authentication_keys = [ :username ]

This is displayed in the rails server console when submitting the form:

Started POST "/users" for ::1 at 2015-03-17 14:29:13 -0700
Processing by Devise::RegistrationsController#create as HTML
  Parameters: {"utf8"=>"✓", "authenticity_token"=>"bc5+ffyBSr6h79aumdMpwHhp5OY69
Tk5oKyY+eIBHLCDDwPmxiMbjbE6OTIaeUPGLmS0J+QlwlGFGHki8SKsgA==", "user"=>{"email"=>
"jessa@example.com", "username"=>"Jessa", "password"=>"[FILTERED]", "password_co
nfirmation"=>"[FILTERED]"}, "commit"=>"Sign up"}
Unpermitted parameter: email
   (0.0ms)  BEGIN
   (0.0ms)  ROLLBACK
  Rendered devise/shared/_links.html.erb (2.0ms)
  Rendered devise/registrations/new.html.erb within layouts/application (11.0ms)

I see Unpermitted parameter: email, but don’t know how to fix it.

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