Personal tip: DO NOT share the keystore with anyone
In addition, you can send an apk message to the client, sign it and send it back. If you do not want to do this, you will need to share the keystore in order to arrive at a friendly solution.
You can try to explain to the customer the security risk that they pose and see if they agree to re-publish this application before even more ratings and downloads are reached.
In addition, you should always create a new keystore for each individual client (or even individual applications).
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